Geophilidae
Geophilus flavus, the Netherlands
Scientific classification
Kingdom:
Phylum:
Class:
Order:
Suborder:
Superfamily:
Family:
Geophilidae

Leach, 1815 [1]
Synonyms[2]

Chilenophilidae
Sogonidae

Geophilus sp. mother and brood.

The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea[3] containing the mostly defunct clades Aphilodontidae,[4] Dignathodontidae,[5] Linotaeniidae,[6] Chilenophilinae,[7] and Macronicophilidae.[8]

Description

Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.[9]

Compared to most other families in the suborder Adesmata, this family features a modest number of leg-bearing segments and limited variation in this number within each species.[10] Two European species in this family include centipedes with only 29 pairs of legs: Geophilus persephones (29 in the only specimen, a male), and G. richardi (29 or 31 in males and 33 in females).[11][12] In the order Geophilomorpha, only two species include centipedes with fewer leg pairs, both of them in the family Schendylidae.[11] Several other species in this family are known from specimens with notably few leg pairs in each sex, including Ribautia platensis (as few as 31 in each sex),[13] G. hadesi (33 in each sex),[14] Schendyloides alacer (as few as 33 in each sex),[15] and Strigamia sibirica (as few as 33 in each sex).[16]

Genera

This family contains these genera:[1]

  • Abatorus
  • Acanthogeophilus
  • Achilophilus
  • Agathothus
  • Agnathodon
  • Algerophilus
  • Alloschizotaenia
  • Aphilodon
  • Apogeophilus
  • Arctogeophilus
  • Arenophilus
  • Aztekophilus
  • Barrophilus
  • Bebekium
  • Bithyniphilus
  • Brachygeophilus
  • Brachygonarea
  • Caliphilus
  • Cephalodolichus
  • Cheiletha
  • Chileana
  • Chilenophilus
  • Chomatophilus
  • Clinopodes
  • Condylona
  • Damothus
  • Dekanphilus
  • Dignathodon
  • Diphyonyx
  • Dyodesmophilus
  • Dysmesus
  • Ecuadoron
  • Endogeophilus
  • Eremerium
  • Eremorus
  • Erithophilus
  • Eurygeophilus
  • Eurytion
  • Fagetophilus
  • Filipponus
  • Galliophilus
  • Garrina
  • Geomerinus
  • Geoperingueyia
  • Geophilus
  • Gnathoribautia
  • Gosipina
  • Hapleurytion
  • Harmostela
  • Harpacticellus
  • Henia
  • Horonia
  • Hovanyx
  • Hyphydrophilus
  • Ketampa
  • Kurdistanius
  • Lionyx
  • Macronicophilus
  • Maoriella
  • Mecistauchenus
  • Mecophilus
  • Mixophilus
  • Nabocodes
  • Nannocrix
  • Navajona
  • Nesidiphilus
  • Nesomerium
  • Nicopus
  • Nothogeophilus
  • Oligna
  • Orinomerium
  • Ortognathus
  • Pachymerellus
  • Pachymerinus
  • Pachymerium
  • Pagotaenia
  • Pandineum
  • Peruphilus
  • Philacroterium
  • Plestophilus
  • Plateurytion
  • Pleurogeophilus
  • Poaphilus
  • Polycricus
  • Polygonarea
  • Porethus
  • Portoricellus
  • Portoricona
  • Proschizotaenia
  • Pseudofagetophilus
  • Purcellinus
  • Pycnona
  • Queenslandophilus
  • Ribautia
  • Schendyloides
  • Schizonampa
  • Schizonium
  • Schizopleres
  • Schizotaenia
  • Sepedonophilus
  • Serrona
  • Sogona
  • Steneurytion
  • Stenotaenia
  • Strigamia
  • Stylolaemus
  • Sundageophilus
  • Synerium
  • Synthophilus
  • Taiyuna
  • Taschkentia
  • Telocricus
  • Timpina
  • Tretechthus
  • Tuoba
  • Tylonyx
  • Watophilus
  • Zantaenia
  • Zygona
  • Zygophilus

References

  1. 1 2 "Geophilidae Leach, 1815". Integrated Taxonomic Information System. Retrieved January 2, 2022.
  2. ↑ "Geophilidae".
  3. ↑ Bonato, Lucio (2014). "Phylogeny of Geophilomorpha (Chilopoda) inferred from new morphological and molecular evidence". Cladistics. 30 (5): 485–507. doi:10.1111/cla.12060. PMID 34794246. S2CID 86204188.
  4. ↑ Calvanese, Victor C. (2019-11-15). "Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily". Zootaxa. 4698 (1): zootaxa.4698.1.1. doi:10.11646/zootaxa.4698.1.1. PMID 32229999. S2CID 209592191. Retrieved 21 December 2021.
  5. ↑ "ITIS - Report: Dignathodontidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  6. ↑ "ITIS - Report: Linotaeniidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  7. ↑ Crabill, Ralph Edwin (1954). "A conspectus of the northeastern North American species of Geophilus (Chilopoda Geophilomorpha Geophilidae)". Proceedings of the Entomological Society of Washington. 56: 172–188. Retrieved 23 October 2021 – via Biodiversity Heritage Library.
  8. ↑ "ITIS - Report: Macronicophilidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  9. ↑ Minelli, Alessandro (2011-03-21). Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda. Brill. pp. 414–424. ISBN 978-90-04-18826-6. Retrieved 24 October 2021.
  10. ↑ Minelli, Alessandro; Bortoletto, Stefano (1988-04-01). "Myriapod metamerism and arthropod segmentation". Biological Journal of the Linnean Society. 33 (4): 323–343. doi:10.1111/j.1095-8312.1988.tb00448.x. ISSN 0024-4066.
  11. 1 2 Pereira, Luis Alberto (2013-01-01). "Discovery of a second geophilomorph species (Myriapoda: Chilopoda) having twenty-seven leg-bearing segments, the lowest number recorded up to the present in the centipede order Geophilomorpha". Papéis Avulsos de Zoologia. 53 (13): 163–185. doi:10.1590/S0031-10492013001300001. hdl:11336/3449. ISSN 1807-0205.
  12. ↑ Bonato, Lucio; Minelli, Alessandro; Drago, Leandro; Pereira, Luis Alberto (2015-09-25). "The phylogenetic position of Dinogeophilus and a new evolutionary framework for the smallest epimorphic centipedes (Chilopoda: Epimorpha)". Contributions to Zoology. 84 (3): 237–253. doi:10.1163/18759866-08403004. hdl:11577/3146565. ISSN 1875-9866.
  13. ↑ Silvestri, Philippus (1898). "Nova Geophiloidea Argentina". Comunicaciones del Museo Nacional de Buenos Aires (in Latin). 1 (2): 39–40 – via Biodiversity Heritage Library.
  14. ↑ Stoev, Pavel; Akkari, Nesrine; Komericki, Ana; Edgecombe, Gregory; Bonato, Lucio (2015). "At the end of the rope: Geophilus hadesi sp. n. – the world's deepest cave-dwelling centipede (Chilopoda, Geophilomorpha, Geophilidae)". ZooKeys (510): 95–114. doi:10.3897/zookeys.510.9614. PMC 4523767. PMID 26257537.
  15. ↑ Pereira, Luis Alberto; Minelli, Alessandro (1992). "A new record of Schendyloides alacer (Pocock, 1891) from the Falkland Islands and a redescription of the species (Chilopoda Geophilomorpha)". Bolletino della Società Entomologica Italiana. 124 (2): 83–90.
  16. ↑ Bonato, Lucio; Danyi, Laszlo; Socci, Antonio Augusto; Minelli, Alessandro (2012-12-20). "Species diversity of Strigamia Gray, 1843 (Chilopoda: Linotaeniidae): a preliminary synthesis". Zootaxa. 3593 (1): 1–39 [8]. doi:10.11646/zootaxa.3593.1.1. ISSN 1175-5334.


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